To quote the docs:
When creating an index, the number associated with a key specifies the direction of the index, so it should always be 1 (ascendin
The simple answer that you are looking for is that the direction only matters when you are sorting on two or more fields.
If you are sorting on {a : 1, b : -1}:
{a : 1, b : -1}
Index {a : 1, b : 1} will be slower than index {a : 1, b : -1}
{a : 1, b : 1}